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  Plantae
      Magnoliophyta
            Magnoliopsida
                    Boraginales
                            Boraginaceae
                                     Actinocarya
                                             Actinocarya tibetica
                                                   [Echinospermum sericeum]
                                                   [Elaeagnus tenuiflora]
                                     Adelocaryum
                                     Aegonychon
                                             Aegonychon zollingeri
                                                   [Buglossoides zollingeri]
                                                   [Lithospermum confertiflorum]
                                                   [Lithospermum japonicum]
                                                   [Lithospermum zollingeri f. albidum]
                                                   [Lithospermum zollingeri]
                                                   [Plagiobothrys zollingeri]
                                     Amblynotus
                                             Amblynotus rupestris
                                                   [Amblynotus obovatus]
                                                   [Eritrichium davuricum]
                                                   [Eritrichium maackii]
                                                   [Eritrichium obovatum]
                                                   [Myosotis davurica]
                                                   [Myosotis obovata]
                                                   [Myosotis rupestris]
                                     Amsinckia
                                             Amsinckia lycopsoides
                                                   [Amsinckia barbata]
                                             Amsinckia menziesii
                                                   [Amsinckia copelandii]
                                                   [Amsinckia eatonii]
                                                   [Amsinckia hirticaulis]
                                                   [Amsinckia hispidissima]
                                                   [Amsinckia micrantha]
                                                   [Amsinckia retrorsa]
                                                   [Amsinckia sparsiflora]
                                                   [Benthamia menziesii]
                                     Anchusa
                                           [Lycopsis]
                                             Anchusa arvensis
                                                   [Anchusa orientalis]
                                                   [Buglossites arvensis]
                                                   [Echioides arvensis]
                                                   [Lycopsis arvensis subsp. orientalis]
                                                   [Lycopsis arvensis]
                                               Anchusa arvensis subsp. orientalis
                                                     [Lycopsis orientalis]
                                     Ancistrocarya
                                             Ancistrocarya japonica
                                                   [Ancistrocarya japonica f. albiflora]
                                     Anoplocaryum
                                             Anoplocaryum compressum
                                             Anoplocaryum tenellum
                                     Antiotrema
                                     Argusia
                                     Arnebia
                                           [Macrotomia]
                                           [Ulugbekia]
                                             Arnebia decumbens
                                                   [Lithospermum cornutum]
                                                   [Lithospermum decumbens]
                                             Arnebia fimbriata
                                             Arnebia guttata
                                                   [Arnebia thomsonii]
                                                   [Arnebia tibetana]
                                                   [Lithospermum guttatum]
                                                   [Macrotomia guttata]
                                     Asperugo
                                             Asperugo procumbens
                                     Beruniella
                                     Borago
                                             Borago officinalis
                                     Borrago
                                     Bothriospermum
                                             Bothriospermum chinense
                                             Bothriospermum kusnetzowii
                                             Bothriospermum secundum
                                                   [Bothriospermum imaii]
                                                   [Bothriospermum secundum f. albiflorum]
                                                   [Lithospermum secundum]
                                             Bothriospermum tenellum
                                             Bothriospermum zeylanicum
                                                   [Anchusa tenella]
                                                   [Anchusa zeylanica]
                                                   [Bothriospermum asperugoides]
                                                   [Bothriospermum perenne]
                                                   [Bothriospermum tenellum var. asperugoides]
                                                   [Cynoglossum prostratum]
                                     Brachybotrys
                                             Brachybotrys paridiformis
                                     Brunnera
                                             Brunnera macrophylla
                                                   [Anchusa myosotidiflora]
                                                   [Myosotis macrophylla]
                                     Buglossites
                                     Buglossoides
                                             Buglossoides arvensis
                                                   [Aegonychon arvense]
                                                   [Lithospermum arvense]
                                                   [Lithospermum sibthoripianum]
                                                   [Rhytispermum arvense]
                                                   [Rhytispermum medium]
                                     Carmona
                                     Casselia
                                     Cerinthe
                                     Cerinthodes
                                     Chionocharis
                                     Coldenia
                                             Coldenia procumbens
                                     Cordia
                                           [Hemigymnia]
                                             Cordia aspera
                                               Cordia aspera subsp. kanehirae
                                                     [Cordia kanehirae]
                                                     [Gerascanthus kanehirae]
                                             Cordia dichotoma
                                     Craniospermum
                                             Craniospermum canescens
                                             Craniospermum mongolicum
                                                   [Craniospermum echioides]
                                             Craniospermum subfloccosum
                                             Craniospermum subvillosum
                                     Cynoglossospermum
                                     Cynoglossum
                                             Cynoglossum alpestre
                                             Cynoglossum amabile
                                             Cynoglossum asperrimum
                                                   [Cynoglossum asperrimum var. tosaense]
                                                   [Cynoglossum asperrimum var. yesoense]
                                                   [Cynoglossum tosaense]
                                                   [Paracynoglossum asperrimum]
                                             Cynoglossum divaricatum
                                             Cynoglossum lanceolatum
                                                   [Cynoglossum formosanum]
                                                   [Cynoglossum lanceolatum var. formosanum]
                                             Cynoglossum officinale
                                                   [Cynoglossum ruderale]
                                                   [Cynoglossum vulgare]
                                             Cynoglossum zeylanicum
                                                   [Cynoglossum formosanum f. albiflorum]
                                                   [Cynoglossum furcatum var. villosulum]
                                                   [Cynoglossum furcatum]
                                                   [Cynoglossum villosulum]
                                                   [Cynoglossum villosum]
                                                   [Cynoglossum zeylanicum f. albiflorum]
                                                   [Cynoglossum zeylanicum var. villosulum]
                                                   [Echinospermum zeylanicum]
                                     Echinospermum
                                     Echioides
                                     Echium
                                             Echium vulgare
                                                   [Echium anglicum]
                                                   [Echium elegans]
                                                   [Echium lacaitae]
                                                   [Echium lusitanicum]
                                                   [Echium molle]
                                                   [Echium monstrosum]
                                                   [Echium schifferi]
                                                   [Echium spinescens]
                                                   [Echium tenoreanum]
                                                   [Echium vaudense]
                                     Ehretia
                                             Ehretia acuminata
                                                   [Cordia thyrsiflora]
                                                   [Ehretia acuminata var. obovata]
                                                   [Ehretia ovalifolia var. latifolia]
                                                   [Ehretia ovalifolia]
                                                   [Ehretia serrata var. obovata]
                                                   [Ehretia taiwaniana]
                                                   [Ehretia thyrsiflora]
                                             Ehretia dicksonii
                                                   [Ehretia dicksonii var. japonica]
                                                   [Ehretia dicksonii var. liukiuensis]
                                                   [Ehretia macrophylla]
                                             Ehretia longiflora
                                             Ehretia microphylla
                                                   [Carmona heterophylla]
                                                   [Carmona microphylla]
                                                   [Carmona retusa]
                                                   [Cordia retusa]
                                                   [Ehretia monopyrena]
                                             Ehretia philippinensis
                                                   [Ehretia dichotoma]
                                                   [Ehretia lanyuensis]
                                                   [Ehretia takaminei]
                                             Ehretia resinosa
                                             Ehretia serrata
                                     Eritrichium
                                             Eritrichium alpinum
                                             Eritrichium borealisinense
                                             Eritrichium canum
                                                   [Echinospermum canum]
                                             Eritrichium fetisowii
                                             Eritrichium incanum
                                                   [Myosotis incana]
                                             Eritrichium jacuticum
                                             Eritrichium kamelinii
                                             Eritrichium mandshuricum
                                             Eritrichium nanum
                                             Eritrichium nipponicum
                                                   [Eritrichium nipponicum f. albiflorum]
                                                   [Eritrichium nipponicum f. yesoense]
                                                   [Eritrichium nipponicum var. albiflorum]
                                                   [Eritrichium nipponicum var. yesoense]
                                                   [Eritrichium sachalinense]
                                                   [Hackelia nipponica]
                                             Eritrichium pauciflorum
                                                   [Eritrichium altaicum]
                                                   [Eritrichium pulviniforme]
                                                   [Eritrichium rupestre]
                                                   [Eritrichium subrupestre]
                                                   [Myosotis pauciflora]
                                             Eritrichium sajanense
                                             Eritrichium sichotense
                                                   [Eritrichium incanum var. sichotense]
                                             Eritrichium spathulatum
                                                   [Echinospermum spathulatum]
                                             Eritrichium thymifolium
                                                   [Echinospermum deflexum var. pumilum]
                                                   [Echinospermum thymifolium]
                                                   [Hackelia thymifolia]
                                                   [Lappula thymifolia]
                                             Eritrichium tianschanicum
                                             Eritrichium villosum
                                                   [Echinospermum sericeum]
                                                   [Eritrichium basifixum]
                                                   [Eritrichium nanum subsp. villosum]
                                                   [Eritrichium sericeum]
                                                   [Myosotis villosa]
                                     Ethretia
                                     Euploca
                                             Euploca procumbens
                                                   [Euploca procumbens var. depressum]
                                                   [Heliotropium gracile var. depressum]
                                                   [Heliotropium ovalifolium var. depressum]
                                                   [Heliotropium procumbens var. depressum]
                                                   [Heliotropium procumbens]
                                     Gastrocotyle
                                     Gerascanthus
                                     Glochidocaryum
                                     Hackelia
                                             Hackelia deflexa
                                                   [Echinospermum deflexum]
                                                   [Echinospermum matsudairae]
                                                   [Eritrichium deflexum]
                                                   [Hackelia matsudairai]
                                                   [Lappula deflexa]
                                                   [Lappula matsudairae]
                                                   [Myosotis deflexa]
                                             Hackelia uncinata
                                                   [Cynoglossum uncinatum]
                                     Heliotropium
                                             Heliotropium arborescens
                                                   [Heliotropium peruvianum]
                                             Heliotropium arboreum
                                                   [Argusia argentea]
                                                   [Heliotropium foertherianum]
                                                   [Messerschmidia argentea]
                                                   [Tournefortia argentea]
                                             Heliotropium eichwaldii
                                             Heliotropium ellipticum
                                             Heliotropium europaeum
                                                   [Alyssum calycinum]
                                                   [Alyssum halimifolium]
                                                   [Alyssum incanum]
                                                   [Arabis halleri]
                                                   [Arabis thaliana]
                                                   [Asperugo procumbens]
                                                   [Brassica campestris]
                                                   [Brassica eruca]
                                                   [Brassica napus]
                                                   [Brassica oleracea var. botrytis]
                                                   [Brassica oleracea]
                                                   [Cheiranthus]
                                                   [Draba alpina]
                                                   [Iberis amara]
                                                   [Sisymbrium murale]
                                                   [Thlaspi arvense]
                                                   [Turritis glabra]
                                             Heliotropium formosanum
                                             Heliotropium gracile
                                             Heliotropium indicum
                                             Heliotropium ovalifolium
                                             Heliotropium sarmentosum
                                                   [Bothriospermum majusculum]
                                                   [Bothriospermum tenellum var. majusculum]
                                                   [Tournefortia sarmentosa]
                                             Heliotropium sibiricum
                                                   [Tournefortia sibirica var. angustior]
                                                   [Tournefortia sibirica]
                                     Henryettana
                                     Heterocaryum
                                     Lacaitaea
                                     Lappula
                                           [Sclerocaryopsis]
                                             Lappula anisacantha
                                                   [Cynoglossospermum anisacanthum]
                                                   [Echinospermum anisacanthum]
                                             Lappula balchaschensis
                                             Lappula brachycentra
                                                   [Echinospermum brachycentrum]
                                             Lappula coronifera
                                             Lappula diploloma
                                             Lappula granulata
                                                   [Lappula gansuensis]
                                                   [Lappula marginata var. granulata]
                                                   [Lappula platyacantha]
                                             Lappula heteracantha
                                                   [Echinospermum heteracanthum]
                                                   [Lappula echinata var. heterantha]
                                                   [Lappula squarrosa subsp. heteracantha]
                                             Lappula intermedia
                                                   [Echinospermum intermedium]
                                             Lappula lipskyi
                                             Lappula macrantha
                                                   [Echinospermum macranthum]
                                             Lappula marginata
                                             Lappula microcarpa
                                                   [Echinospermum microcarpum]
                                                   [Echinospermum oligacanthum]
                                                   [Echinospermum stylosum]
                                                   [Lappula microcarpa var. heterogenea]
                                             Lappula occultata
                                                   [Lappula sinaica var. occultata]
                                             Lappula omphalodes
                                             Lappula patula
                                                   [Echinospermum patulum]
                                                   [Lappula redowskii var. patula]
                                                   [Lappula redowskii]
                                                   [Myosotis redowskii]
                                             Lappula rupestris
                                             Lappula semiglabra
                                                   [Echinospermum semiglabrum]
                                                   [Lappula caspicum]
                                             Lappula sinaica
                                                   [Gerbera connata]
                                                   [Gerbera kunzeana]
                                             Lappula squarrosa
                                                   [Cynoglossospermum consanguineum]
                                                   [Cynoglossospermum lappula]
                                                   [Cynoglossum lappula]
                                                   [Echinospermum consanguineum]
                                                   [Echinospermum cynoglossoides]
                                                   [Echinospermum fabrei]
                                                   [Echinospermum lappula]
                                                   [Echinospermum obtusifolium]
                                                   [Echinospermum squarrosum]
                                                   [Echinospermum vulgare]
                                                   [Lappula consanguinea]
                                                   [Lappula echinata]
                                                   [Lappula myosotis]
                                                   [Myosotis echinata]
                                                   [Myosotis lappula]
                                                   [Myosotis squarrosa]
                                             Lappula stricta
                                                   [Echinospermum strictum]
                                             Lappula tenuis
                                                   [Echinospermum tenue]
                                             Lappula tuvinica
                                     Lasiocaryum
                                           [Oreogenia]
                                     Lepechiniella
                                     Lindelofia
                                             Lindelofia stylosa
                                                   [Cynoglossum stylosum]
                                                   [Lindelofia benthamii]
                                                   [Solenanthus nigricans]
                                     Lithodora
                                     Lithospermum
                                             Lithospermum erythrorhizon
                                                   [Lithospermum officinale subsp. erythrorhizon]
                                                   [Lithospermum officinale var. erythrorhizon]
                                                   [Lithospermum officinale var. japonicum]
                                             Lithospermum hancockianum
                                                   [Nasturtium henryi]
                                                   [Sisymbrium pumilio]
                                             Lithospermum officinale
                                     Maharanga
                                             Maharanga hookeri
                                                   [Onosma hookeri subsp. wardii]
                                     Mattiastrum
                                     Mertensia
                                             Mertensia davurica
                                                   [Casselia davurica]
                                                   [Casselia gracilis]
                                                   [Cerinthodes davuricum]
                                                   [Lithospermum davuricum]
                                                   [Mertensia davurica var. ochroleuca]
                                                   [Mertensia ochroleuca]
                                                   [Pulmonaria amoena]
                                                   [Pulmonaria davurica]
                                                   [Pulmonaria gracilis]
                                                   [Steenhammera davurica]
                                             Mertensia dshagastanica
                                             Mertensia kamczatica
                                             Mertensia maritima
                                                   [Casselia maritima]
                                                   [Casselia parviflora]
                                                   [Mertensia asiatica]
                                                   [Mertensia maritima subsp. asiatica]
                                                   [Mertensia maritima var. asiatica]
                                                   [Pulmonaria procumbens]
                                                   [Steenhammera maritima]
                                             Mertensia pallasii
                                                   [Lithospermum pallasii]
                                             Mertensia pterocarpa
                                                   [Mertensia pterocarpa f. yoshimurae]
                                                   [Mertensia pterocarpa var. yezoensis]
                                                   [Steenhammera pterocarpa]
                                             Mertensia rivularis
                                                   [Lithospermum rivulare]
                                                   [Steenhammera rivularis]
                                             Mertensia sibirica
                                                   [Pulmonaria sibirica]
                                             Mertensia simplicissima
                                                   [Casselia simplicissima]
                                                   [Cerinthodes simplicissimum]
                                                   [Pulmonaria simplicissima]
                                             Mertensia stylosa
                                                   [Cerinthodes stylosum]
                                                   [Pulmonaria stylosa]
                                                   [Steenhammera stylosa]
                                             Mertensia tarbagataica
                                     Metaeritrichium
                                     Microcaryum
                                     Microula
                                           [Schistocaryum]
                                             Microula blepharolepis
                                                   [Omphalodes blepharolepis]
                                             Microula diffusa
                                                   [Omphalodes diffusa]
                                             Microula filicaulis
                                             Microula leucantha
                                             Microula tangutica
                                             Microula tibetica
                                               Microula tibetica var. pratensis
                                                     [Tretocarya pratensis]
                                             Microula trichocarpa
                                                   [Omphalodes trichocarpa]
                                     Myosotis
                                             Myosotis alpestris
                                                   [Myosotis sylvatica f. acaulis]
                                                   [Myosotis sylvatica subsp. alpestris]
                                             Myosotis arvensis
                                                   [Myosotis scorpioides var. scorpioides]
                                             Myosotis austrosibirica
                                             Myosotis baltica
                                             Myosotis caespitosa
                                                   [Myosotis lingulata]
                                                   [Myosotis scorpioides subsp. caespitosa]
                                                   [Myosotis uliginosa]
                                             Myosotis imitata
                                             Myosotis krylovii
                                             Myosotis laxa
                                                   [Myosotis caespitosa var. laxa]
                                                   [Myosotis palustris var. laxa]
                                             Myosotis perennis
                                             Myosotis scorpioides
                                                   [Myosotis palustris]
                                             Myosotis stricta
                                             Myosotis sylvatica
                                                   [Myosotis perennis var. sylvatica]
                                                   [Myosotis sachalinensis]
                                                   [Myosotis sylvatica var. sachalinensis]
                                     Nihon
                                             Nihon akiensis
                                                   [Omphalodes akiensis]
                                             Nihon japonicum
                                                   [Cynoglossum japonicum]
                                                   [Omphalodes japonica f. albiflora]
                                                   [Omphalodes japonica var. echinosperma]
                                                   [Omphalodes japonica]
                                             Nihon krameri
                                                   [Omphalodes krameri f. alba]
                                                   [Omphalodes krameri var. laevisperma]
                                                   [Omphalodes krameri]
                                                   [Omphalodes laevisperma]
                                             Nihon proliferum
                                                   [Omphalodes prolifera]
                                     Nonea
                                             Nonea caspica
                                                   [Lycopsis caspica]
                                                   [Lycopsis picta]
                                                   [Nonea picta]
                                                   [Onosma caspica]
                                             Nonea pulla
                                             Nonea rossica
                                                   [Nonea pulla subsp. rossica]
                                                   [Nonea pulla var. rossica]
                                     Omphalodes
                                     Omphalotrigonotis
                                     Ompholodes
                                     Onosma
                                             Onosma gmelinii
                                             Onosma setosa
                                             Onosma simplicissima
                                             Onosma transrhymnensis
                                                   [Onosma setosa subsp. transrhymnense]
                                     Paracaryum
                                     Paracynoglossum
                                     Pedinogyne
                                     Peristima
                                     Phacelia
                                             Phacelia tanacetifolia
                                                   [Phacelia tanacetifolia var. pseudodistans]
                                     Plagiobothrys
                                     Pulmonaria
                                             Pulmonaria mollissima
                                     Rhytispermum
                                     Rindera
                                           [Mattia]
                                             Rindera tetraspis
                                                   [Cynoglossum laevigatum]
                                                   [Rindera laevigata]
                                     Rochelia
                                             Rochelia bungei
                                                   [Lithospermum retortum]
                                                   [Rochelia disperma]
                                                   [Rochelia retorta]
                                                   [Rochelia stellulata]
                                             Rochelia leiocarpa
                                     Rotula
                                           [Rhabdia]
                                     Sinojohnstonia
                                     Solenanthus
                                             Solenanthus circinnatus
                                                   [Solenanthus coronatus]
                                     Steenhammera
                                     Stenosolenium
                                             Stenosolenium saxatile
                                                   [Anchusa saxatile]
                                                   [Arnebia saxatile]
                                                   [Onosma saxatile]
                                     Strophiostoma
                                     Symphytum
                                             Symphytum officinale
                                                   [Symphytum peregrinum]
                                     Thyrocarpus
                                             Thyrocarpus glochidiatus
                                             Thyrocarpus sampsonii
                                     Tournefortia
                                           [Messerschmidia]
                                     Tretocarya
                                     Trichodesma
                                             Trichodesma calycosum
                                               Trichodesma calycosum var. formosanum
                                     Trigonotis
                                             Trigonotis elevatovenosa
                                                   [Trigonotis formosana var. elevatovenosa]
                                             Trigonotis formosana
                                             Trigonotis guilielmi
                                                   [Eritrichium guilielmi]
                                             Trigonotis icumae
                                                   [Omphalodes icumae]
                                             Trigonotis myosotidea
                                                   [Eritrichium myosotideum]
                                             Trigonotis nankotaizanensis
                                             Trigonotis peduncularis
                                                   [Eritrichium japonicum]
                                                   [Eritrichium pedunculare]
                                                   [Myosotis chinensis]
                                                   [Myosotis peduncularis]
                                                   [Trigonotis clavata]
                                               Trigonotis peduncularis var. amblyosepala
                                             Trigonotis pedunculata
                                             Trigonotis radicans
                                                   [Eritrichium brevipes]
                                                   [Eritrichium radicans]
                                                   [Omphalodes aquatica]
                                                   [Omphalodes sericea]
                                                   [Trigonotis brevipes f. cornata]
                                                   [Trigonotis brevipes var. cornata]
                                                   [Trigonotis brevipes]
                                                   [Trigonotis coreana]
                                                   [Trigonotis cornata]
                                                   [Trigonotis nakaii]
                                                   [Trigonotis radicans subsp. sericea]
                                                   [Trigonotis radicans var. sericea]
                                                   [Trigonotis sericea]
                                     Varronia