Japan Flora: the rhizomes with dark brown hairs; fronds thinner, the lower pinnae broader, lanceolate, about 5 cm. wide; pinnules narrowly ovate to broadly lanceolate-deltoid, 2-3 cm. long, 6-12 mm. wide, obtuse to acute, oblique, short-petiolulate, pinnate to pinnati- parted; ultimate pinnules oblong, toothed; sori nearly marginal, suborbicular; indusia hyaline, glabrous or with few setose hairs near base, undulate to entire.