Northeastern Asian Flora
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Gymnocarpium dryopteris (L.) Newman  
Family: Aspleniaceae
[Aspidium dryopteris (L.) Baumg., moreCarpogymnia dryopteris (L.) ?.L?ve & D.L?ve, Dryopteris dryopteris Christ, Gymnocarpium dryopteris var. aokigaharaense Nakaike, Lastrea dryopteris Bory, Nephrodium dryopteris (L.) Michx., Polypodium dryopteris L., Polypodium dryopteris var. glandulosum Neil, Thelypteris dryopteris (L.) Sloss.]
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Japan Flora: Rhi­

zomes long-creeping, wiry, about 1 mm. across, scaly when young; fronds deciduous, slender, glabrous, often slightly glaucescent on the blades beneath; stipes slender, pale green, lustrous, sparsely scaly, 10-25 cm. long, dark brown at base; scales pale brown, thinly membranous, narrowly ovate, 2-3 mm. long, glabrous, those at base of stipes and on rhizomes broadly ovate, 3-4 mm. long; blades membranous, 5-angled, deltoid, 8-15 cm. long, slightly broader than long, glabrous, smooth; pinnae 3, the terminal as large as to only slightly larger than the lateral ones, deltoid, 5-10 cm. wide, gradually acute, bipinnadparted or in lower part bipinnate, the petioles 1.5-4 cm. long; pinnules deltoid-lanceolate, subacute to obtuse, sessile except for the lowest pair on the terminal pinna, some­times petiolate, the ultimate pinnules narrowly oblong, obtuse, crenate to lobulate; sori in 2 series on the pinnules, nearer the margin than the costule. Coniferous woods.

Hokkaido, Honshu (centr. and n. distr.). Sakhalin, Korea, Himalayas, Siberia to Europe, and N. America.

Gymnocarpium dryopteris
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Development supported by College of Agriculture and Life Sciences of Seoul
National University and Korea National Arboretum of Korea Forest Service.
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