Japan Flora: Rhizomes short-creeping; stipes approximate, 20-40 cm. long, often slightly rubescent; scales on rhizomes and at base of stipes dense, membranous, reddish brown, lanceolate, 8-10 mm. long, to 2 mm. wide, gradually narrowed to a filiform tip, entire, scales on stipes and rachis filiform, deciduous，2-5 mm. long; blades ovate to broadly so, 25-45 cm. long, 15-25 cm. wide, simply pinnate or more often with a pinnate branch on posterior side in the lowest (or lower 2) pairs, the pinnae 2-7 pairs, lanceolate, 12-20 cm. long, 2-3.5 cm. wide, acumi­nate, petiolulate; terminal pinnae acuminate, short-petiolulate; pinnules firmly chartaceous to subcoriaceous, deltoid-ovate to oblong-ovate, 1.5-3 cm. long, 8-12 mm. wide, oblique, short- petiolulate, subacute, spine-tipped and -toothed, slightly lus­trous above, glabrate on both sides; sori submarginal, in 2 ser­ies on the pinnules, nearer the margin than the costule; indusia orbicular-reniform, 1-1.5 mm. across，ciliate, sparsely puberu- lent on back.  

Honshu (Kanto Distr. and westw.), Shikoku, Kyushu.Ryukyus, Formosa, China to Malaysia, and India.

var. yakusimensis (H. Ito) Ohwi. Fronds nar­rower, 45-60 cm. long; pinnae 7-9 pairs, the pinnules smaller, coriaceous, often lobed to parted, 7-15 mm. long; indusia not ciliate. Honshu (Kii Prov.), Shikoku, Kyushu. Ryukyus, Formosa, and China