Korea Flora: Habit: Perennial herb; leaves densely arranged at apex of short rhizome. Stipe: 20-40 cm long, sparsely scaly with blackish-brown scales along with rachis and pinnae axes. Blade: Pentagonal-ovate, broadly ovate; apex abruptly narrowed and acute; 15-20 cm × 15-22 cm; tripinnate, completely or deeply divided. Pinnae: Basal pair largest, broader on posterior side; size abruptly decreasing upwards. Pinnules: Oblong to ovate-oblong; apex acute or obtuse; very shortly stalked; deeply to completely pinnatifid; abaxial surface with bullate scales. Ultimate Segments: Margins with aristate teeth. Sori: Arranged on upper portion of basal veinlets; indusium subreniform with entire margins.
Note: Characterized by pentagonal leaf shape, bullate scales on abaxial surface, and aristate teeth on segments. Habitat: Growing in Baekdu Mountain region.
Japan Flora: Rhizomes short-creeping; stipes 20-50 cm. long, straw-colored, often dark brown toward base, thinly scaly on lower half; scales on stipes and rhizomes pale brown, concolorous, thinly membranous, ovate to broadly so, 3-7 mm. long, acute to abruptly acuminate; blades membranous, deltoid, more or less 5-angled, 15-20 cm. long, 15—22 cm. wide, 3-pinnate or -pin- natiparted, the rachis and secondary axes thinly scaly; pinnae 5-8 pairs, narrowly deltoid, acuminate, distinctly petiolulate, much broader on the posterior than on the anterior side; pinnules oblong-ovate to narrowly oblong, 7-20 mm. long, 2-10 mm. wide, pinnatiparted, obtuse or acute, softly awntoothed, glabrous above, with small ovate scales on veins beneath; sori on upper half of lower veinlets; indusia subreniform, entire, about 0.8 mm. across.Coniferous woods.
Hokkaido, Honshu (n. distr. and n. Kan to Distr.); rare. Sakhalin and Amur.