Korea Flora: Habit: Perennial herb; rhizome short, obliquely ascending; leaves fasciculate. Stipe: 15-30 cm long; densely scaly at base, sparsely scaly above along with rachis. Scales: Brown to dark brown, lanceolate; margins entire. Blade: Pentagonal-broadly-ovate, apex acute; tripinnate; 15-30 cm × 10-22 cm. Pinnae: Elongate-ovate, stalked; basal pair largest. Pinnules: Ovate-oblong; apex obtuse; shortly stalked; 1-2 cm × 4-12 mm; deeply to shallowly pinnatifid; basal pinnules pinnate. Ultimate Segments: Oblong; apex obtuse; margins serrate; 2-3 mm wide; abaxial surface with hair-like scales. Sori: Submarginal; indusium round-reniform with entire margins.
Habitat: Growing in mountainous regions. Note: Characterized by submarginal sori and hair-like scales on abaxial surface of segments.
[Translated from Flora of Korea by T.B. Lee]
Japan Flora: Rhizomes creeping, short, usually densely covered with stubs of old stipes; fronds deciduous; stipes 15-30 cm. long, slender, pale green to brownish, scaly; scales brown, broadly lanceolate, 3-10 mm. long, short-acuminate, remotely toothed to entire; blades 5-angled, broadly ovate, 15-25 cm. long, 10-22 cm. wide, bipinnate, the rachis slender, with spreading scales 1.5-2.5 mm. long; pinnae 5-6 pairs, thinly herbaceous, deltoid-ovate to -lanceolate, 7-15 cm. long, 5-10 cm. wide, acuminate, spreading, the petiolules 1-2.5 cm. long; pinnules ovate-oblong, 1-2 cm. long, 4-12 mm. wide, obtuse, very short-petiolulate, pinnately lobed to parted, or the lowest one pinnatifid; ultimate pinnules oblong, toothed, 2-3 mm. wide, with minute hairlike scales beneath; sori submarginal, near sinus of the teeth; indusia orbicular-reniform, about 0.7 mm. across, entire.
Hokkaido, Honshu, Shikoku, Kyushu. Korea and China.