Japan Flora: Differing chiefly in shorter sori; fronds glabrous, the costas often short-hairy on the underside of the ultimate pinnules; blades ovate-deltoid, bipinnate; pinnules oblong-ovate, 2-4 cm. long, 7-12 mm. wide, acute to obtuse, truncate at base, sessile or very slightly adnate to the scarcely winged rachis, lobulate to parted, obsoletely toothed; sori elliptic, median,about 1 mm. long.